Nodes in a cryptocurrency network are computers responsible for validating transactions and storing blockchain data, which is distinctly different from miners and stakers who rely on node access to the ledger. Nodes include light nodes (for verifying partial data), full nodes (for storing the entire blockchain), and masternodes (for executing governance and supporting additional functions such as private transactions). Masternodes, pioneered by Dash, typically require substantial collateral and resources, while light nodes and full nodes are more accessible and support network decentralization.